“the ground had become impregnated with urine”

[L]ions are not particular about where they urinate. They will do it lying, sitting or standing, at any time and in any place, although they are most particular about where they defecate and will always move well away from their sleeping places. It is quite possible that their free and easy habits of urination have a definite purpose. Lion urine appears to be an insect repellent as I noticed this to be the case of Elsa and her sisters; their small night enclosure became heavily infested by fleas, but after a time, when the ground had become impregnated with urine, the fleas disappeared.

source: A Lifetime with Lions by George Adamson